This is what I was taught if you are using initial caps (or first-letter caps) in your heading:
If the word is three letters and under, use lower case: Four letters and up, use first-letter caps.
For example in this heading it would be: How to use Initial Caps in a Heading
But there are exceptions...
Quick Tip:
Of course the first word in your heading should always have an initial cap no matter how many letters in the word.
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